Fei Chen, Christina Atterby, Per-Henrik Edqvist, Fredrik Pontén, Erik Larsson* and Frank P Ryan. Expression of HERV-R ERV3 encoded Env-protein in human tissues: introducing a novel protein-antibody-based proteomics. J Royal Society of Medicine 2013; 107(1) 22–29.
Villarreal LP and Ryan FP. Viruses in Host Evolution. General Principles and Future Extrapolations; Current Topics in Virology : 79-90.2011; 9
Ryan FP. Human Endogenous Retroviruses in Multiple Sclerosis: Potential for Novel Neuro-Pharmacological Research. Current Neuropharmacology 2011; 9: 360-369.
Ryan FP. Metamorphosis. New Scientist; 24 September 2011: 56-59.
Ryan FP. I, Virus. New Scientist; 30 January 2010: 32-35.
Ryan FP. An alternative approach to medical genetics based on modern evolutionary biology. Part 1: mutation and symbiogenesis. Journal of the Royal Society of Medicine 2009; 102: 272-77.
Ryan FP. An alternative approach to medical genetics based on modern evolutionary biology. Part 2: retroviral symbiosis. Journal of the Royal Society of Medicine 2009; 102: 324-31.
Ryan FP. An alternative approach to medical genetics based on modern evolutionary biology. Part 3: HERVs in diseases. Journal of the Royal Society of Medicine 2009; 102: 415-24.
Ryan FP. An alternative approach to medical genetics based on modern evolutionary biology. Part 4: HERVs in cancer. Journal of the Royal Society of Medicine 2009; 102: 474-80.
Ryan FP. An alternative approach to medical genetics based on modern evolutionary biology. Part 5: epigenetics and genomics. Journal of the Royal Society of Medicine 2009; 102: 530-7.
Ryan FP. Valuable Viruses. Chapter 1 in The Origins of Self, edited by Lynn Margulis, in Press, Chelsea Green Publishers.
Ryan FP. Viruses as symbionts. Symbiosis 2007; 44: 11-21.
Ryan FP (2006). Genomic creativity and natural selection: a modern synthesis. Biological Journal of the Linnean Society 88: 655-72.
Ryan FP (2004). Human endogenous retroviruses in health and disease: a symbiotic perspective. J Royal Soc Medicine.